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A Twitter app that rocks. More wikipedia'ish: A Twitter Application that is so necessary or desirable that it proves the core value of some larger technology (in this case Twitter). A "killer twitter app" can substantially increase sales (adoption in this case) of the platform that it runs on. Ref: http://en.wikipedia.org/wiki/Killer_application
The numbers are accumulated, starting at 0 at the begining of the period.